Saturday, September 29, 2007

The Origins of Indians: Version 9.0

The Dravidians:
Sengupta et al. (2006) makes an interesting comment about spread of Dravidian speakers.

A competing alternative model based on both archaeobotanical material evidence and colloquial agricultural terms, however, more parsimoniously postulates that early Dravidian has a epipaleolithic preagricultural heritage with origins near a South
Asian core region, suggesting possible independent centers of plant domestication within the Indian peninsula by indigenous peoples (Fuller 2003).


They associate Y-Haplogroup L1 with the spread of Dravidian speakers.

Though Y-Haplogroup L1 is present among both Dravidian castes and (most importantly) among Central Dravidian tribes like Koitor (Gond)s, they do not show homogeneous distribution across various agricultural castes in South India.

Also, then there is J2b with a strong presence in South India and which also has rather uniform distribution across castes.

However, R2 dominates agricultural castes (Kamma, Kapu, Reddy) of Telugu region. Also, in certain agricultural castes like Gounders in Tamil Nadu (Sahoo et al. 2006). Except H other haplogroups like, L1, R2 and J2b could be totally absent or present negligibly in some of the south Indian castes. From this point of view only haplogroup H could be associated with the spread of Dravidian languages. Though this can't be ruled out as studies have shown reduced diversity for this haplogroup in India though it is supposedly one of the oldest lineage. This probably can be explained by the scenario where the spread of agriculture increasing its numbers in the last 5000 years. The lineage might have experienced bottleneck in its tribal past.

According to Fuller(2006) agriculture in South India started around 3000BC in south-central Deccan area. I have already mentioned about a paper on spread of Dravidian languages by F C Southworth(University of Pennsylvania). The author proposes a model where Dravidian languages expand from Godavari basin as all the Dravidian branches ND, CD, SD-I,II are observed in that region.

The most important question is whether;
- Dravidian speakers really migrated from north-west of the sub-continent(from Indus valley) or
- was there a language in north-west of India/Pakistan that buffered Dravidian languages of central-south India and various languages of west Asia.

I believe in the latter scenario.

From, population genetics it is very clear that the farthest boundary that can be assigned to Dravidian lineages is east of Indus valley. The distribution of Y-Haplogroups west of Indus valley do not match with Dravidian speakers(H, L1, R2 and J2b).

But there has been speculations that some of the words in Dravidian languages could have been a borrow from west Asian languages (kudure = horse...supposedly a borrow from Elamite language).

Then there is Brahui question. Their presence in west of Indus valley has been taken as a proof for Dravidian speaking past of that region.

However, there are couple of things that I find rather strange about Brahui-s.

- Brahui-s were overwhelmingly herders until 20th century. However, Proto-Dravidian speakers at the best are considered farmers or at the worst pastorals. Therefore, there should have been higher chances that Brahui-s becoming sedentary farmers in a land that developed agriculture around 7000 BCE.

- Brahui-s share R2, L1 and H with Dravidian speakers(totally around 20%). No J2b.

These two exact facts could be used to construct the original urheimat of Dravidian speakers.

Some of the researchers have speculated about independent development of agriculture in south Asia. I would rather consider it as an west Asian extension brought about by J2b clan.

Mehrgarh farming regions around 7000 BCE were developed by J2b clan who later moved south and probably around 3500-3000 BCE brought agricultural knowledge to Dravidian speaking central Indian region. The Dravidian speakers whose major haplogroups were H, L1 and R2 now picked up the knowledge of plant domestication from J2b clan and started expanding through southern region.

However, few herder/pastoral Dravidians who didn't come across J2b clan probably moved to Indo-Iranian regions(Brahui-s as they are known now). Other north Dravidian family speakers like Kurux and Malto are still tribals who migrated to north-eastern regions.

I believe the major agriculture population during Harappan time could have been J2b in the present day Pakistan. Their language could have been closer to Elamite language.

It is said that Elamites did not have their own literary tradition and borrowed everything from their neighbours. Indus valley probably built by the people with the same illiterate cultural background. J2b is a good candidate for that.

Y-Haplogroup association with linguistic expansions:
Austro-Asiatic : O2a
Dravidian : H, R2 and L1
IE : R1a1

Unknown languages:
The language of Indus valley : J2b

Haplogroup association with languages:

Austro-Asiatic : Y-chromosome O2a
Dravidian : mtDNA M
IE : Y-chromosome R1a1

Religious symbol and Language groups or haplogroups:
phallic worship: Austro-Asiatic/Sino-Tibetan
Horse mounted deity: J2b/Dravidian tribes
Brahma and Vishnu: J2a, G2 (Sumerian/Semitic origins)
Spirits/goddesses: tribes/Dravidian tribes
male nature gods: IE speakers

References:

1. The Eurasian Heartland: A continental perspective on Y-chromosome diversity (Wells et al. 2001)
2. A prehistory of Indian Y chromosomes: Evaluating demic diffusion scenarios (Sahoo et al. 2006)
3. Polarity and Temporality of High-Resolution Y-Chromosome Distributions in India Identify Both Indigenous and Exogenous Expansions and Reveal Minor Genetic Influence of Central Asian Pastoralists (Sengupta et al. 2005)
4. Agricultural Origins and Frontiers in South Asia: A Working Synthesis (Fuller 2006)
5. Proto-Dravidian Agriculture (F C Southworth)

Note: All the references are available online.

4 comments:

Anonymous said...

All three/four haplogroups that you find specific of Dravidians are found in Southern Iran - though in smaller proportions - and could mean imports from India, rather than origins, not J2b though. According to Regueiro et al:

- H1a: 2.56% (3/117)
- J2b: 19.65% (23/117) (also: 21.21% in Northern Iran, but less diverse)
- L1: 3.41% (4/117)
- R2: 0.85% (1/117) (also: 3.3%, one hit too, in northern Iran)

Also, you mentioned elsewhere that R2 is more typical of Central-Northern India (the Gujarat-Bengal strip) than of Dravidians.

Couldn't these haplogroups be related with different human waves and their respective cultures? Could, for instance H be Paleolithic and the other haplogroups arrived later?

Anonymous said...

@Maju

-Manju noted earlier that J2a and J2b are switched around. They have J-M410 (normally J2a) listed as J2b.

-H-M69 has Paleolithic duration, also being most regular in Indian tribes. I think it is fairly analogous to I-M170 in Europe in terms of frequency age and distribution. Subclade H1a (most common) seems to have expanded somewhere in India and if it is of Mesolithic age there is a prospect of it belonging to a culture(s) formerly not described.

Ravindra Mundkur said...

Thank you for the specific references. Yet I feel that west coast of India, that is likely to throw more light on migrations. has not been adequately sampled in the genome studies.Southworth assumes that Neolithic cultural data identified in eastern Karnataka (Fuller/Korisettar)represents early Dravidians. I don't think there is any valid basis for this assumption. It might represent the older Ausro-Asiatic or Munda substrata that was subsequently assimilated with Dravidians.
Ibra's suggestion in the comment above seems valid to me.

Manju Edangam said...

Maju, Ibra and Ravi, I'll come back to this topic soon.